// Foundation, Inc., 51 Franklin Street, Fifth Floor, Boston, MA 02110-1301, USA
//
// Contact: phylosoft @ gmail . com
-// WWW: www.phylosoft.org/forester
+// WWW: https://sites.google.com/site/cmzmasek/home/software/forester
package org.forester.sdi;
import java.util.Map;
import java.util.Set;
import java.util.SortedSet;
-import java.util.TreeSet;
import org.forester.phylogeny.Phylogeny;
+import org.forester.phylogeny.PhylogenyMethods;
import org.forester.phylogeny.PhylogenyNode;
import org.forester.phylogeny.data.Event;
-import org.forester.phylogeny.data.Taxonomy;
import org.forester.phylogeny.iterators.PhylogenyNodeIterator;
+import org.forester.sdi.SDIutil.TaxonomyComparisonBase;
import org.forester.util.ForesterUtil;
-/*
- * Implements our algorithm for speciation - duplication inference (SDI). <p>
- * The initialization is accomplished by: </p> <ul> <li>method
- * "linkExtNodesOfG()" of class SDI: setting the links for the external nodes of
- * the gene tree <li>"preorderReID(int)" from class Phylogeny: numbering of
- * nodes of the species tree in preorder <li>the optional stripping of the
- * species tree is accomplished by method "stripTree(Phylogeny,Phylogeny)" of
- * class Phylogeny </ul> <p> The recursion part is accomplished by this class'
- * method "geneTreePostOrderTraversal(PhylogenyNode)". <p> Requires JDK 1.5 or
- * greater.
- *
- * @see SDI#linkNodesOfG()
- *
- * @see Phylogeny#preorderReID(int)
- *
- * @see
- * PhylogenyMethods#taxonomyBasedDeletionOfExternalNodes(Phylogeny,Phylogeny)
- *
- * @see #geneTreePostOrderTraversal(PhylogenyNode)
- *
- * @author Christian M. Zmasek
- */
-public final class GSDI extends SDI {
+public final class GSDI implements GSDII {
- private final boolean _most_parsimonious_duplication_model;
- private final boolean _strip_gene_tree;
- private final boolean _strip_species_tree;
- private int _speciation_or_duplication_events_sum;
- private int _speciations_sum;
- private final List<PhylogenyNode> _stripped_gene_tree_nodes;
- private final List<PhylogenyNode> _stripped_species_tree_nodes;
- private final Set<PhylogenyNode> _mapped_species_tree_nodes;
- private TaxonomyComparisonBase _tax_comp_base;
- private final SortedSet<String> _scientific_names_mapped_to_reduced_specificity;
+ private final boolean _most_parsimonious_duplication_model;
+ private final int _speciation_or_duplication_events_sum;
+ private final int _speciations_sum;
+ private final int _duplications_sum;
+ private final List<PhylogenyNode> _stripped_gene_tree_nodes;
+ private final List<PhylogenyNode> _stripped_species_tree_nodes;
+ private final Set<PhylogenyNode> _mapped_species_tree_nodes;
+ private final TaxonomyComparisonBase _tax_comp_base;
+ private final SortedSet<String> _scientific_names_mapped_to_reduced_specificity;
public GSDI( final Phylogeny gene_tree,
final Phylogeny species_tree,
final boolean most_parsimonious_duplication_model,
final boolean strip_gene_tree,
final boolean strip_species_tree ) throws SDIException {
- super( gene_tree, species_tree );
- _speciation_or_duplication_events_sum = 0;
- _speciations_sum = 0;
+ this( gene_tree, species_tree, most_parsimonious_duplication_model, strip_gene_tree, strip_species_tree, true );
+ }
+
+ public GSDI( final Phylogeny gene_tree,
+ final Phylogeny species_tree,
+ final boolean most_parsimonious_duplication_model,
+ final boolean strip_gene_tree,
+ final boolean strip_species_tree,
+ final boolean transfer_taxonomy ) throws SDIException {
_most_parsimonious_duplication_model = most_parsimonious_duplication_model;
- _duplications_sum = 0;
- _strip_gene_tree = strip_gene_tree;
- _strip_species_tree = strip_species_tree;
- _stripped_gene_tree_nodes = new ArrayList<PhylogenyNode>();
- _stripped_species_tree_nodes = new ArrayList<PhylogenyNode>();
- _mapped_species_tree_nodes = new HashSet<PhylogenyNode>();
- _scientific_names_mapped_to_reduced_specificity = new TreeSet<String>();
- linkNodesOfG();
- getSpeciesTree().preOrderReId();
- geneTreePostOrderTraversal();
+ if ( gene_tree.getRoot().getNumberOfDescendants() == 3 ) {
+ gene_tree.reRoot( gene_tree.getRoot().getChildNode( 2 ) );
+ }
+ final NodesLinkingResult nodes_linking_result = linkNodesOfG( gene_tree,
+ species_tree,
+ strip_gene_tree,
+ strip_species_tree );
+ _stripped_gene_tree_nodes = nodes_linking_result.getStrippedGeneTreeNodes();
+ _stripped_species_tree_nodes = nodes_linking_result.getStrippedSpeciesTreeNodes();
+ _mapped_species_tree_nodes = nodes_linking_result.getMappedSpeciesTreeNodes();
+ _scientific_names_mapped_to_reduced_specificity = nodes_linking_result
+ .getScientificNamesMappedToReducedSpecificity();
+ _tax_comp_base = nodes_linking_result.getTaxCompBase();
+ PhylogenyMethods.preOrderReId( species_tree );
+ final GSDIsummaryResult gsdi_summary_result = geneTreePostOrderTraversal( gene_tree,
+ _most_parsimonious_duplication_model,
+ transfer_taxonomy );
+ _speciation_or_duplication_events_sum = gsdi_summary_result.getSpeciationOrDuplicationEventsSum();
+ _speciations_sum = gsdi_summary_result.getSpeciationsSum();
+ _duplications_sum = gsdi_summary_result.getDuplicationsSum();
}
- GSDI( final Phylogeny gene_tree, final Phylogeny species_tree, final boolean most_parsimonious_duplication_model )
- throws SDIException {
- this( gene_tree, species_tree, most_parsimonious_duplication_model, false, false );
+ public int getDuplicationsSum() {
+ return _duplications_sum;
}
- // s is the node on the species tree g maps to.
- private final void determineEvent( final PhylogenyNode s, final PhylogenyNode g ) {
- boolean oyako = false;
- if ( ( g.getChildNode1().getLink() == s ) || ( g.getChildNode2().getLink() == s ) ) {
- oyako = true;
- }
- if ( g.getLink().getNumberOfDescendants() == 2 ) {
- if ( oyako ) {
- g.getNodeData().setEvent( createDuplicationEvent() );
- }
- else {
- g.getNodeData().setEvent( createSpeciationEvent() );
- }
- }
- else {
- if ( oyako ) {
- final Set<PhylogenyNode> set = new HashSet<PhylogenyNode>();
- for( PhylogenyNode n : g.getChildNode1().getAllExternalDescendants() ) {
- n = n.getLink();
- while ( n.getParent() != s ) {
- n = n.getParent();
- if ( n.isRoot() ) {
- break;
- }
- }
- set.add( n );
- }
- boolean multiple = false;
- for( PhylogenyNode n : g.getChildNode2().getAllExternalDescendants() ) {
- n = n.getLink();
- while ( n.getParent() != s ) {
- n = n.getParent();
- if ( n.isRoot() ) {
- break;
- }
- }
- if ( set.contains( n ) ) {
- multiple = true;
- break;
- }
- }
- if ( multiple ) {
- g.getNodeData().setEvent( createDuplicationEvent() );
- }
- else {
- if ( _most_parsimonious_duplication_model ) {
- g.getNodeData().setEvent( createSpeciationEvent() );
- }
- else {
- g.getNodeData().setEvent( createSingleSpeciationOrDuplicationEvent() );
- }
- }
- }
- else {
- g.getNodeData().setEvent( createSpeciationEvent() );
- }
+ @Override
+ public Set<PhylogenyNode> getMappedExternalSpeciesTreeNodes() {
+ return _mapped_species_tree_nodes;
+ }
+
+ @Override
+ public final SortedSet<String> getReMappedScientificNamesFromGeneTree() {
+ return _scientific_names_mapped_to_reduced_specificity;
+ }
+
+ public final int getSpeciationOrDuplicationEventsSum() {
+ return _speciation_or_duplication_events_sum;
+ }
+
+ @Override
+ public final int getSpeciationsSum() {
+ return _speciations_sum;
+ }
+
+ @Override
+ public List<PhylogenyNode> getStrippedExternalGeneTreeNodes() {
+ return _stripped_gene_tree_nodes;
+ }
+
+ @Override
+ public List<PhylogenyNode> getStrippedSpeciesTreeNodes() {
+ return _stripped_species_tree_nodes;
+ }
+
+ @Override
+ public TaxonomyComparisonBase getTaxCompBase() {
+ return _tax_comp_base;
+ }
+
+ @Override
+ public final String toString() {
+ final StringBuffer sb = new StringBuffer();
+ sb.append( "Most parsimonious duplication model: " + _most_parsimonious_duplication_model );
+ sb.append( ForesterUtil.getLineSeparator() );
+ sb.append( "Speciations sum : " + getSpeciationsSum() );
+ sb.append( ForesterUtil.getLineSeparator() );
+ sb.append( "Duplications sum : " + getDuplicationsSum() );
+ sb.append( ForesterUtil.getLineSeparator() );
+ if ( !_most_parsimonious_duplication_model ) {
+ sb.append( "Speciation or duplications sum : " + getSpeciationOrDuplicationEventsSum() );
+ sb.append( ForesterUtil.getLineSeparator() );
}
+ return sb.toString();
}
/**
* Preconditions: Mapping M for external nodes must have been calculated and
* the species tree must be labeled in preorder.
* <p>
+ * @param transfer_taxonomy
+ * @return
+ * @throws SDIException
*
*/
- final void geneTreePostOrderTraversal() {
- for( final PhylogenyNodeIterator it = getGeneTree().iteratorPostorder(); it.hasNext(); ) {
+ final static GSDIsummaryResult geneTreePostOrderTraversal( final Phylogeny gene_tree,
+ final boolean most_parsimonious_duplication_model,
+ final boolean transfer_taxonomy ) throws SDIException {
+ final GSDIsummaryResult res = new GSDIsummaryResult();
+ for( final PhylogenyNodeIterator it = gene_tree.iteratorPostorder(); it.hasNext(); ) {
final PhylogenyNode g = it.next();
if ( g.isInternal() ) {
+ if ( g.getNumberOfDescendants() != 2 ) {
+ throw new SDIException( "gene tree contains internal node with " + g.getNumberOfDescendants()
+ + " descendents" );
+ }
PhylogenyNode s1 = g.getChildNode1().getLink();
PhylogenyNode s2 = g.getChildNode2().getLink();
while ( s1 != s2 ) {
}
}
g.setLink( s1 );
- determineEvent( s1, g );
+ determineEvent( s1, g, most_parsimonious_duplication_model, res );
+ }
+ if ( transfer_taxonomy ) {
+ transferTaxonomy( g );
}
}
+ return res;
}
- private final Event createDuplicationEvent() {
- final Event event = Event.createSingleDuplicationEvent();
- ++_duplications_sum;
- return event;
- }
-
- private final Event createSingleSpeciationOrDuplicationEvent() {
- final Event event = Event.createSingleSpeciationOrDuplicationEvent();
- ++_speciation_or_duplication_events_sum;
- return event;
- }
-
- private final Event createSpeciationEvent() {
- final Event event = Event.createSingleSpeciationEvent();
- ++_speciations_sum;
- return event;
- }
-
- public final int getSpeciationOrDuplicationEventsSum() {
- return _speciation_or_duplication_events_sum;
+ final static GSDIsummaryResult geneTreePostOrderTraversal( final Phylogeny gene_tree,
+ final boolean most_parsimonious_duplication_model,
+ final int min_duplications ) throws SDIException {
+ final GSDIsummaryResult res = new GSDIsummaryResult();
+ for( final PhylogenyNodeIterator it = gene_tree.iteratorPostorder(); it.hasNext(); ) {
+ final PhylogenyNode g = it.next();
+ if ( g.isInternal() ) {
+ if ( g.getNumberOfDescendants() != 2 ) {
+ throw new SDIException( "gene tree contains internal node with " + g.getNumberOfDescendants()
+ + " descendents" );
+ }
+ PhylogenyNode s1 = g.getChildNode1().getLink();
+ PhylogenyNode s2 = g.getChildNode2().getLink();
+ while ( s1 != s2 ) {
+ if ( s1.getId() > s2.getId() ) {
+ s1 = s1.getParent();
+ }
+ else {
+ s2 = s2.getParent();
+ }
+ }
+ g.setLink( s1 );
+ determineEvent( s1, g, most_parsimonious_duplication_model, res );
+ if ( res.getDuplicationsSum() > min_duplications ) {
+ return null;
+ }
+ }
+ }
+ return res;
}
- public final int getSpeciationsSum() {
- return _speciations_sum;
+ final static NodesLinkingResult linkNodesOfG( final Phylogeny gene_tree,
+ final Phylogeny species_tree,
+ final boolean strip_gene_tree,
+ final boolean strip_species_tree ) throws SDIException {
+ final TaxonomyComparisonBase tax_comp_base = SDIutil.determineTaxonomyComparisonBase( gene_tree );
+ if ( tax_comp_base == null ) {
+ throw new RuntimeException( "failed to establish taxonomy linking base (taxonomy linking base is null)" );
+ }
+ return linkNodesOfG( gene_tree, species_tree, tax_comp_base, strip_gene_tree, strip_species_tree );
}
/**
* This allows for linking of internal nodes of the species tree (as opposed
* to just external nodes, as in the method it overrides.
+ * If TaxonomyComparisonBase is null, it will try to determine it.
* @throws SDIException
*
*/
- @Override
- final void linkNodesOfG() throws SDIException {
+ final static NodesLinkingResult linkNodesOfG( final Phylogeny gene_tree,
+ final Phylogeny species_tree,
+ final TaxonomyComparisonBase tax_comp_base,
+ final boolean strip_gene_tree,
+ final boolean strip_species_tree ) throws SDIException {
+ if ( tax_comp_base == null ) {
+ throw new IllegalArgumentException( "taxonomy linking base is null" );
+ }
final Map<String, PhylogenyNode> species_to_node_map = new HashMap<String, PhylogenyNode>();
final List<PhylogenyNode> species_tree_ext_nodes = new ArrayList<PhylogenyNode>();
- _tax_comp_base = determineTaxonomyComparisonBase( _gene_tree );
+ final NodesLinkingResult res = new NodesLinkingResult();
+ res.setTaxCompBase( tax_comp_base );
// Stringyfied taxonomy is the key, node is the value.
- for( final PhylogenyNodeIterator iter = _species_tree.iteratorExternalForward(); iter.hasNext(); ) {
+ for( final PhylogenyNodeIterator iter = species_tree.iteratorExternalForward(); iter.hasNext(); ) {
final PhylogenyNode s = iter.next();
species_tree_ext_nodes.add( s );
if ( s.getNodeData().isHasTaxonomy() ) {
- final String tax_str = taxonomyToString( s, _tax_comp_base );
+ final String tax_str = SDIutil.taxonomyToString( s, res.getTaxCompBase() );
if ( !ForesterUtil.isEmpty( tax_str ) ) {
if ( species_to_node_map.containsKey( tax_str ) ) {
- throw new SDIException( "taxonomy \"" + s + "\" is not unique in species tree" );
+ throw new SDIException( "taxonomy \"" + tax_str + "\" is not unique in species tree (using "
+ + res.getTaxCompBase() + " for linking to gene tree)" );
}
species_to_node_map.put( tax_str, s );
}
}
}
// Retrieve the reference to the node with a matching stringyfied taxonomy.
- for( final PhylogenyNodeIterator iter = _gene_tree.iteratorExternalForward(); iter.hasNext(); ) {
+ for( final PhylogenyNodeIterator iter = gene_tree.iteratorExternalForward(); iter.hasNext(); ) {
final PhylogenyNode g = iter.next();
if ( !g.getNodeData().isHasTaxonomy() ) {
- if ( _strip_gene_tree ) {
- _stripped_gene_tree_nodes.add( g );
+ if ( strip_gene_tree ) {
+ res.getStrippedGeneTreeNodes().add( g );
}
else {
throw new SDIException( "gene tree node \"" + g + "\" has no taxonomic data" );
}
}
else {
- final String tax_str = taxonomyToString( g, _tax_comp_base );
+ final String tax_str = SDIutil.taxonomyToString( g, res.getTaxCompBase() );
if ( ForesterUtil.isEmpty( tax_str ) ) {
- if ( _strip_gene_tree ) {
- _stripped_gene_tree_nodes.add( g );
+ if ( strip_gene_tree ) {
+ res.getStrippedGeneTreeNodes().add( g );
}
else {
throw new SDIException( "gene tree node \"" + g + "\" has no appropriate taxonomic data" );
}
else {
PhylogenyNode s = species_to_node_map.get( tax_str );
- if ( ( _tax_comp_base == TaxonomyComparisonBase.SCIENTIFIC_NAME ) && ( s == null )
+ if ( ( res.getTaxCompBase() == TaxonomyComparisonBase.SCIENTIFIC_NAME ) && ( s == null )
&& ( ForesterUtil.countChars( tax_str, ' ' ) > 1 ) ) {
- s = tryMapByRemovingOverlySpecificData( species_to_node_map, tax_str );
+ s = tryMapByRemovingOverlySpecificData( species_to_node_map,
+ tax_str,
+ res.getScientificNamesMappedToReducedSpecificity() );
}
if ( s == null ) {
- if ( _strip_gene_tree ) {
- _stripped_gene_tree_nodes.add( g );
+ if ( strip_gene_tree ) {
+ res.getStrippedGeneTreeNodes().add( g );
}
else {
throw new SDIException( "taxonomy \"" + g.getNodeData().getTaxonomy()
}
else {
g.setLink( s );
- _mapped_species_tree_nodes.add( s );
+ res.getMappedSpeciesTreeNodes().add( s );
}
}
}
} // for loop
- if ( _strip_gene_tree ) {
- stripGeneTree();
- if ( getGeneTree().isEmpty() || ( getGeneTree().getNumberOfExternalNodes() < 2 ) ) {
- throw new SDIException( "species could not be mapped between gene tree and species tree" );
+ if ( strip_gene_tree ) {
+ stripTree( gene_tree, res.getStrippedGeneTreeNodes() );
+ if ( gene_tree.isEmpty() || ( gene_tree.getNumberOfExternalNodes() < 2 ) ) {
+ throw new SDIException( "species could not be mapped between gene tree and species tree (based on "
+ + res.getTaxCompBase() + ")" );
}
}
- if ( _strip_species_tree ) {
- stripSpeciesTree( species_tree_ext_nodes );
+ if ( strip_species_tree ) {
+ stripSpeciesTree( species_tree, species_tree_ext_nodes, res );
}
+ return res;
}
- private final PhylogenyNode tryMapByRemovingOverlySpecificData( final Map<String, PhylogenyNode> species_to_node_map,
- final String tax_str ) {
- PhylogenyNode s = tryMapByRemovingOverlySpecificData( species_to_node_map, tax_str, " (" );
- if ( s == null ) {
- if ( ForesterUtil.countChars( tax_str, ' ' ) == 2 ) {
- final String new_tax_str = tax_str.substring( 0, tax_str.lastIndexOf( ' ' ) ).trim();
- s = species_to_node_map.get( new_tax_str );
- if ( s != null ) {
- addScientificNamesMappedToReducedSpecificity( tax_str, new_tax_str );
- }
- }
+ static final void transferTaxonomy( final PhylogenyNode g ) {
+ if ( g == null ) {
+ throw new IllegalArgumentException( "gene tree node is null" );
}
+ final PhylogenyNode s = g.getLink();
if ( s == null ) {
- for( final String t : new String[] { " subspecies ", " strain ", " variety ", " varietas ", " subvariety ",
- " form ", " subform ", " cultivar ", " section ", " subsection " } ) {
- s = tryMapByRemovingOverlySpecificData( species_to_node_map, tax_str, t );
- if ( s != null ) {
- break;
+ throw new IllegalArgumentException( "mapped species tree node is null" );
+ }
+ if ( s.getNodeData().isHasTaxonomy() ) {
+ g.getNodeData().setTaxonomy( s.getNodeData().getTaxonomy() );
+ if ( g.isInternal() ) {
+ if ( g.getChildNode1().isInternal() && g.getChildNode1().getNodeData().isHasTaxonomy()
+ && ( g.getChildNode1().getNodeData().getTaxonomy() == s.getNodeData().getTaxonomy() ) ) {
+ g.getChildNode1().getNodeData().setTaxonomy( null );
+ }
+ if ( g.getChildNode2().isInternal() && g.getChildNode2().getNodeData().isHasTaxonomy()
+ && ( g.getChildNode2().getNodeData().getTaxonomy() == s.getNodeData().getTaxonomy() ) ) {
+ g.getChildNode2().getNodeData().setTaxonomy( null );
}
}
}
- return s;
- }
-
- private final PhylogenyNode tryMapByRemovingOverlySpecificData( final Map<String, PhylogenyNode> species_to_node_map,
- final String tax_str,
- final String term ) {
- final int i = tax_str.indexOf( term );
- if ( i > 4 ) {
- final String new_tax_str = tax_str.substring( 0, i ).trim();
- final PhylogenyNode s = species_to_node_map.get( new_tax_str );
- if ( s != null ) {
- addScientificNamesMappedToReducedSpecificity( tax_str, new_tax_str );
+ else if ( ForesterUtil.isEmpty( g.getName() ) && !ForesterUtil.isEmpty( s.getName() ) ) {
+ g.setName( s.getName() );
+ if ( g.isInternal() ) {
+ if ( g.getChildNode1().isInternal() && ( g.getChildNode1().getName() == s.getName() ) ) {
+ g.getChildNode1().setName( "" );
+ }
+ if ( g.getChildNode2().isInternal() && ( g.getChildNode2().getName() == s.getName() ) ) {
+ g.getChildNode2().setName( "" );
+ }
}
- return s;
}
- return null;
}
- private final void addScientificNamesMappedToReducedSpecificity( final String s1, final String s2 ) {
- _scientific_names_mapped_to_reduced_specificity.add( s1 + " -> " + s2 );
+ private final static void addScientificNamesMappedToReducedSpecificity( final String s1,
+ final String s2,
+ final SortedSet<String> scientific_names_mapped_to_reduced_specificity ) {
+ scientific_names_mapped_to_reduced_specificity.add( s1 + " -> " + s2 );
}
- public final SortedSet<String> getReMappedScientificNamesFromGeneTree() {
- return _scientific_names_mapped_to_reduced_specificity;
- }
-
- public TaxonomyComparisonBase getTaxCompBase() {
- return _tax_comp_base;
- }
-
- private void stripSpeciesTree( final List<PhylogenyNode> species_tree_ext_nodes ) {
- for( final PhylogenyNode s : species_tree_ext_nodes ) {
- if ( !_mapped_species_tree_nodes.contains( s ) ) {
- _species_tree.deleteSubtree( s, true );
- _stripped_species_tree_nodes.add( s );
- }
+ private final static void determineEvent( final PhylogenyNode s,
+ final PhylogenyNode g,
+ final boolean most_parsimonious_duplication_model,
+ final GSDIsummaryResult res ) {
+ boolean oyako = false;
+ if ( ( g.getChildNode1().getLink() == s ) || ( g.getChildNode2().getLink() == s ) ) {
+ oyako = true;
}
- _species_tree.clearHashIdToNodeMap();
- _species_tree.externalNodesHaveChanged();
- }
-
- public List<PhylogenyNode> getStrippedSpeciesTreeNodes() {
- return _stripped_species_tree_nodes;
- }
-
- private void stripGeneTree() {
- for( final PhylogenyNode g : _stripped_gene_tree_nodes ) {
- _gene_tree.deleteSubtree( g, true );
+ if ( g.getLink().getNumberOfDescendants() == 2 ) {
+ if ( oyako ) {
+ g.getNodeData().setEvent( Event.createSingleDuplicationEvent() );
+ res.increaseDuplicationsSum();
+ }
+ else {
+ g.getNodeData().setEvent( Event.createSingleSpeciationEvent() );
+ res.increaseSpeciationsSum();
+ }
}
- _gene_tree.clearHashIdToNodeMap();
- _gene_tree.externalNodesHaveChanged();
- }
-
- public Set<PhylogenyNode> getMappedExternalSpeciesTreeNodes() {
- return _mapped_species_tree_nodes;
- }
-
- public static TaxonomyComparisonBase determineTaxonomyComparisonBase( final Phylogeny gene_tree ) {
- int with_id_count = 0;
- int with_code_count = 0;
- int with_sn_count = 0;
- int max = 0;
- for( final PhylogenyNodeIterator iter = gene_tree.iteratorExternalForward(); iter.hasNext(); ) {
- final PhylogenyNode g = iter.next();
- if ( g.getNodeData().isHasTaxonomy() ) {
- final Taxonomy tax = g.getNodeData().getTaxonomy();
- if ( ( tax.getIdentifier() != null ) && !ForesterUtil.isEmpty( tax.getIdentifier().getValue() ) ) {
- if ( ++with_id_count > max ) {
- max = with_id_count;
+ else {
+ if ( oyako ) {
+ final Set<PhylogenyNode> set = new HashSet<PhylogenyNode>();
+ for( PhylogenyNode n : g.getChildNode1().getAllExternalDescendants() ) {
+ n = n.getLink();
+ while ( ( n.getParent() != s ) && ( n.getParent() != null ) ) {
+ n = n.getParent();
+ if ( n.isRoot() ) {
+ break;
+ }
}
+ set.add( n );
}
- if ( !ForesterUtil.isEmpty( tax.getTaxonomyCode() ) ) {
- if ( ++with_code_count > max ) {
- max = with_code_count;
+ boolean multiple = false;
+ for( PhylogenyNode n : g.getChildNode2().getAllExternalDescendants() ) {
+ n = n.getLink();
+ while ( ( n.getParent() != s ) && ( n.getParent() != null ) ) {
+ n = n.getParent();
+ if ( n.isRoot() ) {
+ break;
+ }
}
+ if ( set.contains( n ) ) {
+ multiple = true;
+ break;
+ }
+ }
+ if ( multiple ) {
+ g.getNodeData().setEvent( Event.createSingleDuplicationEvent() );
+ res.increaseDuplicationsSum();
}
- if ( !ForesterUtil.isEmpty( tax.getScientificName() ) ) {
- if ( ++with_sn_count > max ) {
- max = with_sn_count;
+ else {
+ if ( most_parsimonious_duplication_model ) {
+ g.getNodeData().setEvent( Event.createSingleSpeciationEvent() );
+ res.increaseSpeciationsSum();
+ }
+ else {
+ g.getNodeData().setEvent( Event.createSingleSpeciationOrDuplicationEvent() );
+ res.increaseSpeciationOrDuplicationEventsSum();
}
}
}
+ else {
+ g.getNodeData().setEvent( Event.createSingleSpeciationEvent() );
+ res.increaseSpeciationsSum();
+ }
}
- if ( max == 0 ) {
- throw new IllegalArgumentException( "gene tree has no taxonomic data" );
- }
- else if ( max == 1 ) {
- throw new IllegalArgumentException( "gene tree has only one node with taxonomic data" );
- }
- else if ( max == with_id_count ) {
- return SDI.TaxonomyComparisonBase.ID;
- }
- else if ( max == with_sn_count ) {
- return SDI.TaxonomyComparisonBase.SCIENTIFIC_NAME;
+ }
+
+ private final static void stripSpeciesTree( final Phylogeny species_tree,
+ final List<PhylogenyNode> species_tree_ext_nodes,
+ final NodesLinkingResult res ) {
+ for( final PhylogenyNode s : species_tree_ext_nodes ) {
+ if ( !res.getMappedSpeciesTreeNodes().contains( s ) ) {
+ species_tree.deleteSubtree( s, true );
+ res.getStrippedSpeciesTreeNodes().add( s );
+ }
}
- else {
- return SDI.TaxonomyComparisonBase.CODE;
+ species_tree.clearHashIdToNodeMap();
+ species_tree.externalNodesHaveChanged();
+ }
+
+ private final static void stripTree( final Phylogeny phy, final List<PhylogenyNode> strip_nodes ) {
+ for( final PhylogenyNode g : strip_nodes ) {
+ phy.deleteSubtree( g, true );
}
+ phy.clearHashIdToNodeMap();
+ phy.externalNodesHaveChanged();
}
- public List<PhylogenyNode> getStrippedExternalGeneTreeNodes() {
- return _stripped_gene_tree_nodes;
+ private final static PhylogenyNode tryMapByRemovingOverlySpecificData( final Map<String, PhylogenyNode> species_to_node_map,
+ final String tax_str,
+ final SortedSet<String> scientific_names_mapped_to_reduced_specificity ) {
+ PhylogenyNode s = tryMapByRemovingOverlySpecificData( species_to_node_map,
+ tax_str,
+ " (",
+ scientific_names_mapped_to_reduced_specificity );
+ if ( s == null ) {
+ if ( ForesterUtil.countChars( tax_str, ' ' ) == 2 ) {
+ final String new_tax_str = tax_str.substring( 0, tax_str.lastIndexOf( ' ' ) ).trim();
+ s = species_to_node_map.get( new_tax_str );
+ if ( s != null ) {
+ addScientificNamesMappedToReducedSpecificity( tax_str,
+ new_tax_str,
+ scientific_names_mapped_to_reduced_specificity );
+ }
+ }
+ }
+ if ( s == null ) {
+ for( final String t : new String[] { " subspecies ", " strain ", " variety ", " varietas ", " subvariety ",
+ " form ", " subform ", " cultivar ", " section ", " subsection " } ) {
+ s = tryMapByRemovingOverlySpecificData( species_to_node_map,
+ tax_str,
+ t,
+ scientific_names_mapped_to_reduced_specificity );
+ if ( s != null ) {
+ break;
+ }
+ }
+ }
+ return s;
}
- @Override
- public final String toString() {
- final StringBuffer sb = new StringBuffer();
- sb.append( "Most parsimonious duplication model: " + _most_parsimonious_duplication_model );
- sb.append( ForesterUtil.getLineSeparator() );
- sb.append( "Speciations sum : " + getSpeciationsSum() );
- sb.append( ForesterUtil.getLineSeparator() );
- sb.append( "Duplications sum : " + getDuplicationsSum() );
- sb.append( ForesterUtil.getLineSeparator() );
- if ( !_most_parsimonious_duplication_model ) {
- sb.append( "Speciation or duplications sum : " + getSpeciationOrDuplicationEventsSum() );
- sb.append( ForesterUtil.getLineSeparator() );
+ private final static PhylogenyNode tryMapByRemovingOverlySpecificData( final Map<String, PhylogenyNode> species_to_node_map,
+ final String tax_str,
+ final String term,
+ final SortedSet<String> scientific_names_mapped_to_reduced_specificity ) {
+ final int i = tax_str.indexOf( term );
+ if ( i > 4 ) {
+ final String new_tax_str = tax_str.substring( 0, i ).trim();
+ final PhylogenyNode s = species_to_node_map.get( new_tax_str );
+ if ( s != null ) {
+ addScientificNamesMappedToReducedSpecificity( tax_str,
+ new_tax_str,
+ scientific_names_mapped_to_reduced_specificity );
+ }
+ return s;
}
- sb.append( "mapping cost L : " + computeMappingCostL() );
- return sb.toString();
+ return null;
}
}